The prehistoric Croods family live in a particularly dangerous moment in time. Patriarch Grug, his mate Ugga, teenage daughter Eep, son Thunk, and feisty Gran gather food by day and huddle together in a cave at night. When a more evolved caveman named Guy arrives on the scene, Grug is distrustful, but it soon becomes apparent that Guy is correct about the impending destruction of their world.

Searching for a safer habitat, the prehistoric Crood family discovers an idyllic, walled-in paradise that meets all of its needs. Unfortunately, they must also learn to live with the Bettermans -- a family that's a couple of steps above the Croods on the evolutionary ladder. As tensions between the new neighbors start to rise, a new threat soon propels both clans on an epic adventure that forces them to embrace their differences, draw strength from one another, and survive together.