A womanizing city man meets Tillie in the country. When he sees that her father has a very large bankroll for his workers, he persuades her to elope with him.

Tillie Todd gives her rich Aunt Sally a dime-store tomato pincushion for her birthday, claiming that it is the product of her own hard labor. However, Cousin Percy, also competing for Sally's favor, snidely exposes the deception. While testing his latest invention, Tillie's paramour, The Bat, makes a crash landing at Aunt Sally's birthday celebration. Inspired, eccentric Sally takes to the air herself, but after several days' absence she is declared dead. Aunt Sally bequeaths Tillie the tomato pincushion, giving most of the estate to her pet monkey, Jim, and after Jim's death, to Percy. Wanting to speed up his inheritance, Percy kidnaps Jim and connives to frame Tillie for his apparent death. The police relentlessly pursue Tillie until Aunt Sally finally returns. Disgusted at her inability to rid herself of the ugly pincushion, which Percy nastily keeps returning to her, Tillie rips it apart with her teeth, only to discover that it is filled with 10,000 dollar bills.

Tillie and her neighbor Mr. Pipkins are both distraught over their respective marriages. One day, they sneak off to have a lively time at Coney Island. They flee the park together just as their spouses come to find them. After a chase, each is rescued from the ocean and reconcile with their respective spouses.